- Dynamics of dissolved and particulate organic matter in river systems

- Metabolism (production and respiration)

- The microbial compartment and its connection with other compartments

2-. Arrival and dynamics of nutrients and organic materials

3.- Flux and arrival of particulate organic matter.

4.- Structure and function of biofilms

5.- Metabolism, in relation to the arrival of materials and its use by heterotrophs.

Practical

1-. Collection of hydrological, chemical and physiographical data in a stream.

2.- Activities of primary producers and heterotrophs in a stream. Laboratory and field approaches.